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Hamsher, S. E.; Graeff, C. L.; Stepanek, J. G.; Kociolek, J. P. (2014). Frustular morphology and polyphyly in freshwater Denticula (Bacillariophyceae) species, and the description of Tetralunata gen. nov. (Epithemiaceae, Rhodopalodiales). Plant Ecology and Evolution. 147(3): 346-365.